etau—these two, namely Rāma and Kṛṣṇa; hi—certainly; viśvasya—of the universe; ca—and; bīja-yonī—both the cause and ingredient; rāmaḥ—Balarāma; mukundaḥ—Kṛṣṇa; puruṣaḥ—the original Mahā-Viṣṇu; pradhānam—material energy; anvīya—after entering; bhūteṣu—into the material elements; vilakṣaṇasya—of varieties of manifestation; jñānasya—of knowledge; ca—also; īśāte—are the controlling power; imau—both of Them; purāṇau—are the original cause.